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Fossils of Ontario

Part 2: Macroinvertebrates and

Vertebrates of the Champlain Sea

with a listing of nonmarine species

Frances J. E. Wagner

io Museum Life Sciences Miscellaneous Publications

Digsiti^ed by the Internet Archive

iff 2012 with funding from

Royal Ontario Museum

http://archive.org/details/fossilsofontario02bolt

LIFE SCIENCES MISCELLANEOUS PUBLICATIONS
ROYAL ONTARIO MUSEUM

Frances J. E. Wagner Fossils of Ontario

Part 2:

Macroinvertebrates
and Vertebrates of the
Champlain Sea

with a listing of
nonmarine species

<£

ROM

ROYAL ONTARIO MUSEUM
PUBLICATIONS IN LIFE SCIENCES

The Royal Ontario Museum publishes three series in the Life Sciences:

Contributions: a numbered series of original scientific publications.

Occasional Papers: a numbered series of original scientific publications, primarily short and of

taxonomic significance.

Miscellaneous Publications: an unnumbered series on a variety of subjects.

All manuscripts considered for publication are subject to the scrutiny and editorial'policies of
the Life Sciences Editorial Board, and to independent refereeing by two or more persons, other
than Museum staff, who are authorities in the particular field involved.

LIFE SCIENCES EDITORIAL BOARD

Senior editor: J. R. Tamsitt

Editor: D. R. Calder

Editor: J. C. Barlow

External editor: C. S. Churcher

Manuscript editor: P. H. von Bitter

Production editor: J. E. Hawken

Frances J. E. Wagner is a palaeontologist formerly with the Environmental Marine Geology
Subdivision of the Atlantic Geoscience Centre, a Division of the Geological Survey of Canada
based at Dartmouth, Nova Scotia.

Canadian Cataloguing in Publication Data

Main entry under title:
Fossils of Ontario

(Life sciences miscellaneous publications, ISSN

0082-5093)

Contents: v. 1. The trilobites / Rolf Ludvigsen — v. 2.

Macroinvertebrates and vertebrates of the Champlain

Sea / Frances J. E. Wagner.

Bibliography: p.

ISBN 0-88854-221-6 (v. 1). - ISBN 0-88854-295-X (v. 2)

1. Paleontology — Ontario. I. Ludvigsen, Rolf. 1944-
The trilobites. II. Wagner, Frances J. E., 1927-
Macroinvertebrates and vertebrates of the Champlain
Sea. III. Royal Ontario Museum. IV. Series.

QE748.06F67 560.9713 C83-098578-6

Cover illustration: Ophiura sarsi Liitken, impression in clay, p. 36.

Publication date: 29 June 1984

ISBN 0-88854-295-X

ISSN 0082-5093

100 Queen's Park, Toronto, Canada, M5S 2C6

PRINTED AND BOUND IN CANADA AT THE ALGER PRESS

Contents

Acknowledgements 5
Introduction 7

Champlain Sea Deposits in Ontario 9
History of Champlain Sea Studies in Ontario
Champlain Sea Fossils 10
Invertebrata 12
Porifera 1 2

Tethya 1 3
Bryozoa 13

Porella 13
Gastropoda 13

Margarites 1 4

Velutina 1 5

Lunatia 1 5

Natica 1 6

Neptunea 1 7

Admete 17

Cvlichna 17

P/u'/me 19

Haminoea 1 9
Pelecypoda 19

Nucula 1 9

Nuculana 23

Portlandia 23

Portlandia sensu stricto 24

Portlandia (Yoldiella) 24

Mytilus 25

Thyasira 26

Axinopsida 26

/Istarte 26

Macoma 27

A/va 27

Hiatella 28
Conchostraca 30

Cyzicus 30
Cirripedia 30

Balanus 3 1

Balanus sensu stricto 31

Balanus (Chirona) 3 1
Isopoda 33

Mesidotea 33
Asteroidea 33

Crossaster 33
Ophiuroidea 34

Ophiocoma 37

Ophiura 37
Polychaeta 37
Nereis 37
Serpula 37
Vertebrata 39
Chordata 39
Pisces 39

Clupeiformes 39
Mallotus 39
Osmerus 40
Gadiformes 40

Microgadus 40
Gasterosteiformes 40

Gasterosteus 42
Perciformes 42
Artediellus 42
Cyclopterus 42
Mammalia 43
Cetacea 44
Delphinapterus 44
Megaptera AA
Balaena 44
Pinnipedia 44
Phoca AA
Erignathus Al
Nonmarine Species 50
Insecta 50
Pisces 50
Mammalia 50
Aves 50
Plantae 51
Repositories of Illustrated Specimens 53
Glossary of Morphological Terms 54
Literature Cited 59

Acknowledgements

I am most grateful to T. E. Bolton, Geological Survey of Canada (GSC), and C. R.
Harington, National Museums of Canada, for their efforts in procuring photographs
that enabled me to fill the gaps in my coverage of species' illustrations. The
illustrations, exclusive of sketches, are the work of the photographic units of the GSC
and the National Museums of Canada (NMC), both in Ottawa, and of the Bedford
Institute of Oceanography (810), Dartmouth, Nova Scotia. I am indebted to the staff
of the drafting and illustrations office of the BIO for the surficial geology map and for
the sketches of specimens.

My thanks go also to C. T. Schafer, Atlantic Geoscience Centre (AGC), and to
T. E. Bolton, again, for critical reading of the manuscript.

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Introduction

The last continental glaciation in North America, the Wisconsinan, saw the major
part of Canada covered by an extensive ice sheet. In eastern Canada the weight of the
ice depressed the surface of the land by at least several hundred metres, and with the
retreat of the glaciers, marine waters flowed inland to cover much of the Ottawa-St.
Lawrence Lowland. The part of this marine inundation lying west of Quebec City has
been named the Champlain Sea. In eastern Ontario dated marine sediments indicate
that the waters of this sea extended up the Ottawa Valley to the vicinity of
Rapides-des-Joachims, about 65 km upriver from Pembroke, and west at least as far
as Brockville in the St. Lawrence Valley (Fig. 1).

During the Champlain Sea's geologically brief history, from about 12 500 years
B.P. to 9 500 years B.P., various types of animals, ranging from microscopic
foraminifera to whales, penetrated from the Atlantic Ocean into this inland sea. Their
remains have been found embedded in the marine sediments of the area. This book
emphasizes the groups of animals generally large enough to be visible to the naked
eye; in other words, it excludes the microscopic foraminifera and ostracods, which
are to be dealt with in a future volume of "Fossils of Ontario".

The types of animals that lived in the Champlain Sea and that are covered in this
volume comprise sponges, bryozoans (moss animals), molluscs (gastropods and
pelecypods, known otherwise as snails and clams), arthropods (a conchostracan or
clam shrimp, barnacles, and an isopod, which is related to the terrestrial wood lice),
an asteroid (starfish), ophiuroids (brittlestars), annelid worms, fish, whales, and
seals. With the exception of one genus of barnacle, two genera of pelecypods, and
one genus of seal, each genus is represented by only a single species. Nonmarine
fossils found in concretions at Green Creek (Green's Creek of early literature), in the
Ottawa area, are listed but not described.

Molluscs are the most common both in variety and numbers. White gastropod and
pelecypod shells are easily visible in a variety of sediments. A branchiopod, several
genera of fish, and the bones of a young seal are other marine fossils that have been
found in the Green Creek concretions. These concretions offer very attractive and
interesting specimens. Bones of whales and seals are of rare occurrence in the
Champlain Sea deposits.

Champlain Sea fossil occurrences have been known since the late 1830s and their
assemblages have been reported in various scientific journals and government
publications. I recorded the Champlain Sea fossil references dating from 1837 in a
Geological Survey of Canada paper (Wagner, 1967). I have abstracted the references
pertinent to the Ontario scene from this and from other papers published after 1966.
Earlier papers are often vague as to localities of fossil discoveries, citing occurrences
such as "Ottawa and vicinity" or "right bank of Rideau River, near Manotick
Road". More recent references are usually more specific and can lead the searcher to
the exact sites where the fossils have been collected. New localities are continually
being discovered, and a visit to such places as newly dug ditches, foundation
excavations, fresh landslide scars, actively worked clay, and sand and gravel pits can
reward the collector with interesting suites of fossils.

Species found in the Champlain Sea still exist, and those interested in the
fascinating exercise of attempting to determine past environmental conditions and
changes in these conditions can do so by comparing the fossil assemblages with their
modern counterparts. Pelecypod fossils with their valves intact and in living position
are abundant locally, as are apparently in situ gastropod shells. The specimens so
preserved are those of species generally favouring a fine-grained substrate, and they
are found in the clays and silts. The more widespread sands and gravels, being
shallow-water deposits, were subject to wave action which, of course, affected the
animal hard parts enclosed in them. Disarticulated pelecypod valves and barnacle
plates, eroded gastropod shells, and fragments of all kinds are the common fossils in
this type of deposit although complete specimens may be found. Fossils in the
concretions are usually well preserved, but their occurrence is very localized.
Detailed species descriptions and information pertaining to environmental factors
favourable to the various species may be found in Abbott (1974), Bousfield (1960),
Collins (1959), and Morris (1973).

Illustrations of complete specimens are the ideal for identification purposes
because they give a whole against which fragmentary specimens can be compared.
Unfortunately, many Ontario-collected specimens are incomplete, and therefore,
specimens from the province are in the minority among the following figures. In
addition, many specimens of the species reported in older literature could not be
located, and sometimes even specimens from more recent collections were found to
be missing. Where necessary, specimens (either existent or fossil) from elsewhere
have been substituted or have been shown in addition to less than ideal Ontario
material. For a few species, no substitute specimens could be obtained, and so
drawings have been prepared.

Champlain Sea Deposits in Ontario

Unconsolidated deposits in the Ontario area of the Champlain Sea show a sequence of
sediments of glacial, through freshwater, to brackish and marine, and back to
freshwater origin. The brackish and marine materials relate to the Champlain Sea
inundation. Sediment distributions shown in Figure 1 are based primarily on my own
field observations. Fine-grained silts and clays are characteristic of the earlier, deeper
phases of the sea, whereas sands and gravels were deposited during shallower
intervals and along the shores. Boulder beaches were developed where till ridges
projecting above the level of the marine waters allowed the waves to winnow out the
fine-grained sediments.

The earlier names of Leda clay and Saxicava sand have been replaced by the more
appropriate Champlain Sea clay and Champlain Sea sand (Gadd, 1960). Leda is a
synonym of Nuculana; the pelecypod species characteristic of the clays is a
Portlandia, namely Portlandia arctica (Gray). Saxicava is a synonym of Hiatella.

References for description and discussion of the Champlain Sea stratigraphy are
given below.

GENERAL REFERENCES SPECIFIC REFERENCES

Antevs(1925) Ami (1887, 1892, 1906)

Chalmers (1907) Ells (1898, 1907)

Coleman (1901a, 1901b, 1932, 1941) Gadd (1963a, 1963b, 1977, 1980)
Dawson, J. W. (1871, 1883b, 1893) Johnston (1917)
Logan (1863) Keele and Johnston (1913)

Murray (1852) Kindle (1918)

Owen (1951)
Richard (1975)
Terasmae (1960, 1965)
Wilson, W. J. (1898)

History of Champlain Sea Studies in Ontario

Observations of fossiliferous Champlain Sea sand and clay in Quebec (Beauport area)
date back to 1837 when shells were collected there by H. W. Bayfield (1837). They
were identified by Sir Charles Lyell (1841). The first record of Champlain Sea fossils
in Ontario dates to 1 845 when Lyell reported the occurrence of the capelin Mallotus
villosus. The fish-bearing concretions had been obtained by William (later Sir
William) Logan, founder and first director of the Geological Survey of Canada. They
came from the shores of the Ottawa River near Ottawa (then called Bytown). Logan
published subsequent reports on fossil finds from the Ottawa area and from north and
west of Ottawa as far as Lake Coulonge, the major contribution appearing in the 1863
"Geology of Canada". Two other collectors of the 1850s who advanced the
knowledge of the composition and distribution of Champlain Sea faunas in Ontario

were Alexander Murray of the Geological Survey of Canada and Joseph Leidy, an
American palaeontologist.

An important collector in the period 1857 to 1895 was J. William (later Sir
William) Dawson. In his later years he was principal of McGill College in Montreal.
Dawson added considerably to the number of species known from the Champlain Sea
and in 1893, he published the definitive work up to that time on the Pleistocene of
Canada. His son, George Mercer Dawson, was third director of the Geological
Survey of Canada, and his report on the operations of the Geological Survey for 1895
(published in 1897) makes reference to N. J. Giroux's collection of Champlain Sea
fossils from the St. Lawrence area.

During the latter part of the 19th century and the first decade of the 20th Henry M.
Ami published extensively on the Champlain Sea. He added significantly to the lists
of both invertebrate and vertebrate species and recorded new localities, mainly in and
around Ottawa. Ami was a palaeontologist with the Geological Survey at the time.

The 20th century saw the proliferation of people interested in the Champlain Sea
and its faunas. Some of them merely repeated what was already known, sometimes
adding a species or two or a new locality; others undertook detailed studies of a
palaeoecologic or a stratigraphic nature. Major contributors of palaeoecologic
interpretations include Winnifred Goldring of the State Museum at Albany, New
York, and E. J. Whittaker and F. J. E. Wagner, the latter two invertebrate
palaeontologists with the Geological Survey of Canada. Whittaker's observations
were based on collections from along the St. Lawrence River between Prescott,
Ontario and Lachine, Quebec. Goldring considered material from the Ottawa and
Montreal areas although her main emphasis was on the area around Lake Champlain
in Vermont and New York State. She studied the relationship of size differences in
selected species to ecologic conditions, primarily to salinity. The full extent of the
Champlain Sea was considered by Wagner, whose interpretation of ecologic
conditions complemented and extended that of Goldring. She also added materially to
the number of known fossiliferous sites in Ontario and Quebec.

Other contributors since the tum of the century to the knowledge of the Champlain
Sea species and their distribution in Ontario include Ernst Antevs, Robert Bell,
Robert Chalmers, R. W. Ells, W. A. Johnston, E. M. Kindle, Lawrence M. Lambe,
C. M. Sternberg, Jaan Terasmae, and J. F. Whiteaves, all of whom were associated
with the Geological Survey of Canada. Persons other than staff of the Geological
Survey whose works on the Champlain Sea have been cited in the list of references
include A. P. Coleman, C. R. Harington, J. W. Laverdiere, G. H. Perkins, and
H. G. Richards. All references relate to locality citations and not to taxonomic
works.

Champlain Sea Fossils

Excluding the species of foraminifera and ostracods, at least 47 marine species have
been identified from the area of the Champlain Sea in Ontario as opposed to a total of
approximately 125 species from the Champlain Sea as a whole. No new marine
species in addition to the 47 were collected from the parts of Quebec adjacent to
Ontario. It is necessary to go as far as Montreal and south and east from there to find
species other than those reported from Ontario.

The 47 marine species from Ontario are distributed among 40 genera of which 29
are invertebrates. A breakdown of the invertebrates shows one genus of Porifera
(sponges), of Bryozoa (moss animals), of Conchostraca (clam shrimps), of Cirripedia
(barnacles), of Isopoda, and of Asteroidea (starfish); two genera of Ophiuroidea
(brittlestars) and of Polychaeta (worms); nine of Gastropoda (snails); and ten of
Pelecypoda (clams). Among the vertebrates there are five genera of mammals and six
genera of fish.

Organisms found in the Champlain Sea deposits may be classified as follows. The
taxonomic categories under which the various forms are described appear in boldface.

PHYLUM

SUPERCLASS

CLASS

SUBCLASS

ORDER

Porifera

Demospongia

Epipolasida

Bryozoa

Mollusca

Gastropoda
Pelecypoda

Arthropoda

Crustacea

Branchiopoda
Cirripedia

Malacostraca

Conchostraca
Isopoda

Echinodermata

Stelleroidea

Asteroidea
Ophiuroidea

Annelida

Polychaeta

Chordata

Pisces

Osteichthyes
Mammalia

Actinopterygii

Clupeiformes

Gadiformes

Gasterosteiformes

Perciformes

Cetacea

Pinnipedia

Invertebrata

Porifera (Sponges) Figure 2A-D

Porifera are represented by only one genus, Tethya, a sponge belonging to the class
Demospongia Solles, 1875, and the order Epipolasida Solles, 1888. These are
sponges characterized by a complex structure of many chambers. The spicules may
be either siliceous or of spongin, a flexible material related to horn. Spicules of both
kinds may be present. Spicules are of two types, namely megascleres and
microscleres (Fig. 2A-C). Megascleres make up the skeletal framework of the
sponge, whereas microscleres are present in the flesh and do not form part of the
framework. Microscleres are very rarely found in fossils. Epipolasids are typically
radiate in form with a relatively thick external cover. The main spicules are strongyles
(Fig. 2A) with swollen, spindlelike shafts.

Ami (1897, 1902) referred the single species from the Champlain Sea to the genus
Craniella and Whittaker (1922) placed Craniella as a subgenus of Tethya (spelled

Fig. 2 Porifera (A-D) Bryozoa (E)
A Megasclere (a strongyle).
B Microsclere (a euaster).
C Microsclere (a sphaeraster).
Tethya logani Dawson

D Flattened rosettes of spicules, x 3.25, GSC 20130, Montreal, Quebec.
Porella elegantula d'Orbigny
E Schematic representation of several zooecia, X 2.5.

Tethea in earlier publications). However, the two genera are distinct (Moore, 1955,
p. E42). The specimen identified asCraniella cranium (Miiller) by Whiteaves (1901)
is probably Tethya logani.

Tethya Lamarck Figure 2D

This is the one genus found in the Champlain Sea. Tethya is of spheroidal form with a
warty, leathery surface. Megascleres in this genus are strongyles (Fig. 2A) and
microscleres include sphaerasters and euasters (Fig. 2B, C). As fossils, only the
megascleres have been reported. These spicules are found in clay as isolated
individuals, in bundles resembling mouse fur, or clustered as flattened rosettes. The
spicules are needlelike, hollow, gently curved, and tapering at both ends. Some are
drawn out to very fine points, whereas others are more bluntly rounded. The
megascleres measure up to 7.5 mm in length.

A specimen from the Montreal area of Quebec is illustrated.
SPECIES Tethya logani Dawson [recorded as Craniella logani, Tethea logani,

Tethea {Craniella) logani]
REFERENCES Ami (1887, 1892, 1897, 1902), Wagner (1970), Whittaker (1922)

Bryozoa (Moss Animals) Figure 2E

Remains of these colonial animals are very scarce in the Champlain Sea deposits.
Only one genus, Porella, which appears in the earlier literature asEschara, has been
identified positively from the Ottawa area. Fragments of bryozoan zoaria were
reported from Stormont County but were not identifiable. The colonies may be free,
branching forms or they may be encrusting on stones or shells. The zoaria can be seen
readily with the naked eye but a hand lens is needed in order to see the details of the
zooecia.

Porella Gray Figure 2E

Porella is a foliate form. The apertural face may either be smooth or have a
granulated calcareous covering over the smooth inner layer. There is a suboral
avicularium. The ovicell is imperforate.

No specimen was available for illustration.
SPECIES Porella elegantula d'Orbigny
REFERENCES Ami (1892, 1897, 1902), Goldring (1922)

Gastropoda (Snails) Figures 3 to 5

Gastropods, commonly known as snails, are molluscs with a single calcareous shell
that is closed at the apex and is not divided into regular chambers. Shell shapes and
ornamentation are diverse. In most genera the shell is asymmetric.

Gastropods are one of the more common fossils in the Champlain Sea sediments,
although their variety is smaller and their numbers far fewer than those of pelecypods.
With the exception of Neptunea, all are small, ranging in length from 6 mm to about
25 mm. Neptunea may reach 120 mm in length. Many living gastropods have
coloured shells, some of which are quite elaborately patterned. However, the genera
represented here are dull, for the most part, ranging from amber to tan to brown and

ADAPICAL

ADAXIAL

ABAXIAL

CALLUS
UMBILICUS

OPERCULUM

Fig. 3 Morphological terms applied to gastropods.

RETICULATE

SPIRAL STRIAE

SHOULDER
RIB (COSTA)

SUTURE

PARIETAL
REGION

INNER LIP

-OUTER LIP

APERTURE

COLUMELLA

ANTERIOR
(SI PHONAL)
CANAL

brownish grey in life. Some show colour banding. Fossil specimens are almost
invariably bleached white, although occasionally they show a trace of colour or a
colour pattern.

Shells may be found in clays and silts and in sands and gravels, but the best source
is the former group, which are the finer-grained sediment. Most specimens reported
during the period between 1897 and the present have been from the Ottawa area. This
may reflect merely the fact that a growing city with its many excavations offered
excellent chances for the discovery of these shells. However, my field program in the
1950s, which covered all of the area of Ontario that had been inundated by the
Champlain Sea waters, brought to light extremely few samples of gastropods outside
the Ottawa area. Their distribution is probably related to ecology. Most gastropod
species identified favour a clay, silt, or fine sand substrate, and these sediments are
best developed and exposed in the Ottawa Valley.

Figure 3 illustrates features of shell morphology and Figure 4 shows various
gastropod shapes.

Margarites Gray Figure 5A

This is a small gastropod, generally less than 15 mm in length. It is trochiform in
shape and has a rounded aperture with an interrupted peristome. The usually small,
deep umbilicus is encircled by a spiral cord. A distinctive feature of the genus is the

Fig. 4 Some common gastropod shapes.
A Cylindrical. C Globose.
B Fusiform. D Trochiform.

E Subglobose.

nacreous interior of the aperture. The exterior of the shell may be smooth or spirally
ribbed.

Two species have been identified from the Montreal area, but the single damaged
specimen from Ottawa could not be identified as to species. Because the Ottawa fossil
specimen was missing, a modern specimen representing one of the species reported
from Quebec has been illustrated.
SPECIES Margarites sp.
REFERENCE Wagner (1970)

Velutina Fleming Figure 5B

Velutina is small, ranging up to about 20 mm in length. The shell is very thin and
subglobose with two or three rapidly enlarging whorls that may be spirally striated or
reticulately ornamented. The outer lip of the aperture is thin.

Finds in Ontario have been in and around Ottawa; the genus is more commonly
represented in Quebec. A modern specimen is shown in place of the missing original
fossil.

SPECIES Velutina undata Brown
REFERENCES Ami (1902), Whiteaves (1901)

Lunatia Gray Figure 5C

This is another gastropod scarce in Ontario, although it has been reported from
several localities in Quebec. Shells of the genus may attain a length of 125 mm.
However, the species found in the Champlain Sea does not grow more than 25 mm
long. The globose shell has steadily enlarging, rounded whorls, and there is a small
umbilicus, which may be almost closed. The aperture is long, between two-thirds and

three-quarters of the total length of the shell, and wide, but of variable proportions.

A specimen from Ontario was not available, and therefore a specimen from Quebec
is illustrated.

SPECIES Lunatia pallida (Broderip and Sowerby)
REFERENCE Goldring (1922)

Natica Scopoli Figure 5D

Natica is similar in shape to Lunatia but is smaller (maximum length for the genus
about 32 mm) and has its umbilicus almost or completely sealed with a callus. If the
operculum is present, its composition is an important diagnostic feature; the
operculum of Natica is calcareous, whereas that of Lunatia is horny.

Specimens have been collected from several localities in and around Ottawa and
also from Farran Point, a site now inundated by the waters of the St. Lawrence
Seaway. Like Lunatia, Natica is more widely recorded in Quebec. No specimen from
Ontario could be found for illustration.

Fig. 5 Margarites Velutina Lunatia Natica

Margarites costalis (Gould)

A A modern specimen, x 2, GSC 21096, Hudson Bay.

Velutina undata Brown

B A modern specimen, x 12, GSC 55158, Baie des Chaleurs.

Lunatia pallida (Broderip and Sowerby)

C Oral view, x 2, GSC 20136, near Ste-Genevieve, Quebec.

Natica clausa Broderip and Sowerby

D Oral view, X 2, GSC 20135, near Ste-Genevieve, Quebec.

SPECIES Natica clausa Broderip and Sowerby £V. affinis Gmelin is a synonym]
references Ami (1897, 1902), Antevs (1925, 1939), Goldring (1922), Johnston
(1916, 1917), Whittaker (1922)

Neptunea Roding Figure 6A

Representatives of this genus are the largest gastropods one is likely to find in the
Champlain Sea deposits. Specimens from these deposits may reach 75 mm in length;
the largest species of the genus reach about 125 mm. The fusiform shells generally
have six to eight whorls. Sculpture is mostly spiral on the rounded or prominently
shouldered whorls. The spiral ribs may be of two sizes. Some specimens have
indistinct axial ribs. Sutures between adjoining whorls may be deep or shallow. The
aperture is characterized by a short to long, usually twisted, siphonal canal.

In Ontario, this genus is apparently confined to the Ottawa district. A specimen
from Quebec has been shown in place of the unavailable Ontario material.
SPECIES Neptunea despecta (Linne)
references Ami (1902), Antevs (1925, 1939), Goldring (1922), Johnston (1916,

1917)

Ad mete Kroyer Figure 6B

Admete has a thin but strong shell about 25 mm long. The aperture is equal to about
half the total length of the shell, which is basically fusiform in shape. The rounded
whorls are sculptured with strong axial ridges and weak spiral threads. The aperture
has a short, open, barely perceptible anterior canal. There are weak folds on the
columella.

This gastropod is rare; it has been reported only once in Ontario, from Ottawa.
Elsewhere in the Champlain Sea deposits, it has been collected from localities in
Montreal and Quebec City. The whereabouts of Goldring's specimen is unknown to
me; a sketch is presented because no suitable specimen of this species was available
to be photographed.

SPECIES Admete couthouyi (Jay) [A. viridula (Fabricius) is a synonym]
reference Goldring (1922)

Cylichna Loven Figure 6C

This and the following two genera are "bubble" shells and range from globose to
cylindrical in shape. Cylichna is characterized by a small (9 mm or less in length),
cylindrical shell that is generally smooth and glossy, although some species are
ornamented with microscopic spiral scratches. The spire is involute, resulting in a
small apical concavity. The lower end of the shell is truncate. The aperture is long
and narrow, expanded below, and with a single oblique fold on the columella.

Only one of four species identified from the Champlain Sea has been found in
Ontario. Several localities in and around Ottawa have been cited. A single specimen,
which could not be identified to the species level, came from the Cornwall area. This
gastropod has been collected from both sands and clays. A modern individual has
been illustrated because no fossil material could be located.
SPECIES Cylichna alba (Brown)
references Ami (1897, 1902), Antevs (1925, 1939), Goldring (1922), Johnston

(1916, 1917), Wagner (1958)

Fig. 6 Neptunea Admete Cylichna Philine Haminoea

Neptunea despecta (Linne)

A Oral view of specimen previously referred to N. despecta tornata (Gould),

GSC 20140, Grande-Riviere-du-Chene, Quebec.
Admete couthouyi (Jay)
B Schematic representation, x 2.
Cylichna alba (Brown)

C Oral view of a modern specimen, x 6, GSC 55112, Bay of Fundy.
Philine lima (Brown)
D Schematic representation, x 15.
Haminoea solitaria (Say)
E Oral view, x 7, GSC 20141, Apple Hill, Ontario.

x 1,

Philine Ascanius Figure 6D

This is the smallest of the gastropods that one is likely to find, ranging from 2 to
7 mm in length. The shell is thin and loosely coiled with a flaring aperture. In some
species the top of the aperture is above the apex of the shell; in others it is below. The
apex of the shell may be partly flattened or it may be rounded. Ornamentation is
usually spiral and may consist of finely scalloped lines, or rows of small oblong rings,
or microscopic punctations.

There is only one record of this genus in Ontario — near Ottawa. The only other
record of the genus in the Champlain Sea deposits is from the Montreal area. In the
absence of Goldring's specimen and suitable modern material, a drawing is
presented.

SPECIES Philine lima (Brown) [P. lineolata Couthouy is a synonym]
REFERENCE Goldring (1922)

Haminoea Turton and Kingston Figure 6E

This "bubble" shell is similar in shape to the others, Cylichna and Philine, being
globose to broadly cylindrical. The shell of Haminoea, however, may be partly
corneous and is often semitransparent and flexible. The apex of the shell is perforate;
a diagnostic feature for species of this genus is the direction in which the lip of the
aperture rises — to the right or to the left of the perforation, when viewed with the
apex of the shell towards you and with the outer lip facing to the right. The sides of
the whorls may be flattened or rounded. Ornamentation may be lacking or may
consist of spiral grooves.

This genus was recorded earlier from the Ottawa area and from Kenyon Twp.,
Glengarry County.
SPECIES Haminoea solitaria (Say)
references Goldring (1922), Wagner (1970)

Pelecypoda (Clams) Figures 7 to 15

Pelecypods are two-shelled organisms with wholly or partly calcified valves covering
the right and left sides of the body. The hinge plate, with the distinctive pattern and
shape of its teeth, is of diagnostic importance at the generic level. In some genera the
hinge is edentulous. A dorsal ligament connects the valves that open and close along
an anteroposterior axis. The surface of the valves may be smooth, with only faint
concentric growth lines, or it may be highly sculptured concentrically, radially, or
both. General features of shell morphology are shown in Figure 7.

Pelecypods are the most abundant macro fossils in the Champlain Sea deposits.
Complete shells of infaunal species may be found in living position (Fig. 8), and it is
not uncommon to see an exposure, particularly a gravel pit, liberally sprinkled with
white pelecypod valves (Fig. 9). Separated valves may be concentrated, convex side
up, by wave or current action. Living positions of several typical genera are shown in
Figure 10.

Nucula Lamarck Figure HA

The valves of this small (less than 9 mm long), ovate-to-trigonal pelecypod close
tightly all around the periphery. The outer surface of the shell is either smooth or

DORSAL

UMBO

BEAK
CARDINAL TOOTH
HINGE PLATE

LIGAMENT AREA

ADDUCTOR

MUSCLE

SCAR

VENTRAL

POSTERIOR ADDUCTOR
MUSCLE SCAR

PALLIAL SINUS

PALLIAL LINE
CRENULATE MARGIN

LUNULE
GROWTH LINE

CHONDROPHORE

LIGAMENT

LU
>

_l
<
>

I-
I

RESILIFER
Fig. 7 Morphological terms applied to pelecypods.

POSTERIOR

ornamented with fine radial and concentric striae, and the interior of the shell is
pearly. Interiorly, the ventral margins may be smooth or crenulate. Species identified
from the Champlain Sea belong to the subgenus Leionucula , which has smooth inner
ventral margins. A resilifer separates the anterior and posterior sections of the hinge
with its closely interlocking, taxodont teeth. Beaks in this genus are opisthogyrate.
There is no pallial sinus.

Fig. 8 Pelecypod shells (Hiatella arctica) in living position (above point of hammer), gravel
pit 4.6 km south of Alexandria, Ontario (GSC Photo 144026).

Representatives of this genus have been found in the clays at various localities in
the Ottawa Valley. Only one species has been identified, and it is apparently more
common in Ontario than elsewhere. However, because the original material could not
be found, a specimen from the Champlain Sea deposits in Quebec is shown.
SPECIES Nacula tenuis Montagu; Nucula sp. [the N. expansa listed by Coleman

from Pakenham (?) is probably N. tenuis]
REFERENCES Ami (1902), Antevs (1925, 1939), Coleman (1901b), J. W. Dawson

Fig. 9 Exposure liberally sprinkled with pelecypod shells (foreground), gravel pit 4.6 km
south of Alexandria, Ontario (GSC Photo 144027).

HIATELLA

MYTILUS

MY A

Fig. 10 Living positions of selected pelecypod genera.

(1857, 1871, 1893), Goldring (1922), Johnston (1916), Kindle (1918), Logan
(1863), Whiteaves (1901), Whittaker (1922)

Nuculana Link Figure 11B, C

Nuculana has an elongate, generally rostrate shell, either smooth or with concentric
sculpture and with a polished periostracum. The interior of the shell is porcellaneous
and the ventral margins are always smooth. Shells range from about 6 to 38 mm in
length. The chevron-shaped hinge teeth are in two series separated by a wide,
posteriorly directed resilifer. There are approximately twice as many teeth in the
posterior series as in the anterior. There is a small pallial sinus.

Only one of the species of Nuculana recorded from the Champlain Sea has been
found in Ontario and it is of limited occurrence. It has been reported from the Ottawa
area and from near Farran Point, one of the communities obliterated by the St.
Lawrence Seaway. No fossil material was available for illustration so a modern
specimen has been substituted and the interior view has been sketched.
SPECIES Nuculana minuta (Fabricius)
REFERENCES Goldring (1922), Whittaker (1922)

Portlandia (Morch) Figure 12

Portlandia and Nuculana belong to the same family, Nuculanidae; however,

Fig. 1 1 Nucula Nuculana

Nucula tenuis Montagu

A Interior and exterior views (part of hinge has been broken), x 4, GSC 20144,

Lachevrotiere River, Quebec.
Nuculana minuta (Fabricius)

B Exterior of left valve, x 2, GSC 22037, Hudson Bay.
C Diagram to show typical dentition.

Portlandia differs from the latter genus in that the surfaces of its valves are
unornamented and in that the posterior end of its ovate-to-elliptical shell is dorsally
produced. The interior of the shell of Portlandia, as of Nuculana, is porcellaneous,
and the margins are smooth. A large resilifer separates the two series of
chevron-shaped teeth. The number of teeth is approximately the same in both series,
although there may be slightly fewer in the anterior series. If a pallial sinus is present,
it is small. Specimens may reach 30 mm in length.

Portlandia sensu stricto Figure 12 A, B

The typical Portlandia has a well-developed, albeit small, pallial sinus. Its resilifer is
large and subtriangular. The maximum length is 30 mm, but the length of most
specimens is only about 15 mm. The one species in the Champlain Sea, Portlandia
(Portlandia) arctica , is ubiquitous in the marine clays and is often found in silts and
silty sands as well. It is the Leda glacialis, Leda arctica, or Yoldia arctica of the
earlier authors and is the shell for which the Leda clay was named. Actually, Leda is a
synonym of Nuculana, not of Portlandia.
SPECIES Portlandia {Portlandia) arctica (Gray)

references Ami (1884, 1897, 1902), Antevs (1925, 1939), Coleman (1901b,
1941), J. W. Dawson (1857, 1871, 1893), Goldring (1922), Johnston (1916,
1917), Kindle (1918), Lowdon and Blake (1973, 1979), Lowdon, Fyles, and Blake
(1967), Richards (1962), Terasmae (1960, 1965), Wagner (1958, 1970),
Whiteaves(1901), Whittaker (1922), A. E. Wilson (1956), W. J. Wilson (1898)

Portlandia (Yoldiella) Verrill and Bush Figure 12C

Species of this subgenus are small, from 2 or 3 mm to about 15 mm in length, with
most less than 10 mm. The pallial sinus is very small and indistinct, or absent. The

Fig. 12 Portlandia (Portlandia) Portlandia (Yoldiella)

Portlandia (Portlandia) arctica (Gray)

A Interior of left valve, x 4, GSC 551 16, Ottawa, Ontario.

B Exterior of right valve, x 4, GSC 551 17, Ottawa, Ontario.

Portlandia (Yoldiella) lenticula (Moller)

C Interior and exterior views, modern specimen, x 6.75, GSC 551 15. Beaufort Sea.

only species, P. (Y.) lenticula, is recorded from Green Creek, near Ottawa. Leda
pygmaea of earlier authors is possibly this species. It has been necessary to use a
modern specimen for illustration.
SPECIES Portlandia {Yoldiella) lenticula (Moller)

REFERENCES Richards (1962); for Leda pygmaea — Coleman (1901b), J. W.
Dawson (1857, 1871, 1893), Goldring (1922), Logan (1863)

Mytilus Linne Figure 13A

The shell is roughly wedge-shaped with the beaks forming a pointed apex and with
the other end wider and rounded. The colour of the shell is distinctive, being bluish
black on the exterior, but pearly white on the interior, with a deep purple-blue
margin. The surface may be smooth or may have radial ribs. There is a shiny

Fig. 13 Mytilus Thyasira Axinopsida Astarte

Mytilus edulis Linne

A Interior and exterior views of a worn specimen, x 2, GSC 20149, St-Joseph-du-

Lac, Quebec.
Thyasira flexuosa (Montagu)

B Schematic representation of right and left valves, x 1.75.
Axinopsida orbiculata (Sars)

C Interior and exterior views, x 4, GSC 20154, Cornwall area, Ontario.
Astarte montagui (Dillwyn)
D Interior and exterior views of specimen previously referred to A. montagui

striata (Leach), x 2, GSC 20151, St-Janvier-de-Joly, Quebec.

periostracum. Four to six small, weak teeth are located close to the beaks. Some
species may attain a length of 250 mm, but the species encountered in the Champlain
Sea ranges in length between about 25 and 75 mm.

Mytilus is widely distributed and is found mainly in coarser sediments (pebbles to
cobbles and boulders) because it is an attached form. Although complete specimens
may be found, most occurrences are of fragments on!-y. Two tiny pearls (0.35 mm
and 0.4 mm in diameter) were found with well-preserved Mytilus shells in a gravel pit
east of Winchester. Unfortunately, the good shells from Winchester were missing, as
were the older collections. Therefore a specimen from Quebec has been shown.
SPECIES Mytilus edulis Linne

references Ami (1884, 1892, 1897, 1902), Antevs (1925, 1939), Coleman
(1901a, 1901b, 1941), Dawson (1857), Goldring (1922), Johnston (1917), Kindle
(1918), Leidy (1856), Lowdon and Blake (1979), Terasmae (1960, 1965), Wagner
(1958, 1970), Whiteaves (1901), Whittaker (1922), A. E. Wilson (1956)

Thyasira Lamarck Figure 13B

Most representatives of this genus are small, usually less than 15 mm in length. The
posterior area of the subglobular-to-oblique shell is characterized by one or more
radial furrows. The weak hinge is edentulous, although there may be a small
pseudocardinal tubercle in the left valve. The pallial line is without a sinus.
Externally the shell may be chalky or polished and is possibly ornamented with
concentric growth lines but otherwise smooth.

Only a single species has been collected from Ottawa, and because no specimens
were available to be photographed a sketch has been substituted.
SPECIES Thyasira flexuosa (Montagu)
REFERENCES Antevs (1925, 1939)

Axinopsida Keen and Chavan Figure 13C

This genus is similar in shape to Thyasira, but is smaller, being not more than 5 mm
long. The two genera differ also in that Axinopsida does not have a posterior radial
furrow, or furrows, and has a deeply impressed, concave lunule, a feature lacking in
Thyasira. The shell surface is smooth and shiny, sometimes with incremental lines,
and with a thin periostracum. Interiorly, the shell is porcellaneous and polished; the
margins are smooth. The hinge is without teeth, although there may be a thickening
below the beak in each valve forming a peglike pseudocardinal. There is no sinus in
the weakly impressed pallial line.

The Cornwall area has yielded the only representative of this genus so far identified
from the Champlain Sea.
SPECIES Axinopsida orbiculata (Sars)
reference Wagner (1970)

Astarte Sowerby Figure 13D

The shape of this genus may be quadrate, trigonal, or rounded. The surface may be
smooth but more commonly has fine-to-coarse, rounded, concentric ridges. The thick
periostracum ranges from yellow through shades of brown to almost black. Interior
margins of the shell may be smooth or crenulated; in the subgenus recorded from the
Champlain Sea they are smooth. Hinge teeth are variable in shape, development, and
number, but there are always three in the left valve. The pallial line is without a sinus.

Some species of Astarte attain a length of at least 50 mm, A. montagui, found in the
Champlain Sea deposits in Ontario, reaches a maximum length of about 20 mm.

One of two species of Astarte from the Champlain Sea has been found in Ontario,
primarily in and around Ottawa. Earlier authors recorded it as Astarte compressa, A.
banksi , or A. laurentiana. Varieties have been named on the basis of height to
lengthen ratios; however, since the ratios intergrade, it is better to disregard these
varieties.

A specimen from Quebec is illustrated in place of the unavailable material from
Ontario.

SPECIES Astarte montagui (Dillwyn)
REFERENCES Antevs (1925, 1939), Goldring (1922), Johnston (1916, 1917)

Macoma Leach Figure 14 A -D

One species of Macoma reaches a length of more than 100 mm, but the two species
found in the Champlain Sea attain a maximum length of about 50 mm and are usually
much smaller than that. Shells are ovate to subtrigonal, with a posterior flexure that is
usually to the right. The exterior of the shell is generally white, porcellaneous to
chalky, and smooth or with faint concentric growth lines. The hinge line has cardinal
teeth only, two in each valve. A characteristic feature is the configuration of the
pallial sinus, which is of a different size and shape in each valve.

One species, Macoma balthica, is particularly abundant throughout the area; the
other, M. calcarea, is less common but still of widespread occurrence. Both are
generally associated with the sand and gravels, although a few specimens have come
from the clays of the area.

SPECIES Macoma balthica (Linne) [listed as Tellina groenlandica, Macoma
groenlandica, or Macoma fragilis in the earlier literature]; Macoma calcarea
(Gmelin)
references Ami (1884, 1892, 1897, 1902, 1906), Antevs (1925, 1939), Bell
(1906), Chalmers (1907), Coleman (1901a, 1901b, 1941), G. M. Dawson (1897),
J. W. Dawson (1871, 1893), Dyck and Fyles (1963, 1964), Goldring (1922),
Johnston (1916, 1917), Keele and Johnston (1913), Kindle (1918), Leidy (1856),
Logan (1863), Lowdon and Blake (1970, 1973, 1979), Richards (1962), Terasmae
(1960, 1965), Wagner (1958, 1970), Whiteaves (1901), Whittaker (1922), A. E.
Wilson (1956), W. J. Wilson (1898)

Mya Linne Figure 15A, B

This is one of the largest pelecypods to inhabit the Champlain Sea, with specimens as
long as 150 mm possible although individuals of 50 to 75 mm in length are more
usual. The chalky shell is ovate to elongate in shape, rounded anteriorly, and either
produced or truncated posteriorly. The right valve is slightly larger than the left, the
valves closing with either an anterior or a posterior gape. Sculpture is lacking, but
there may be irregular growth lines. The edentulous hinge has a large, projecting,
spoon-shaped chondrophore in the left valve and a recessed ligamental cavity in the
right valve. There is a well-developed pallial sinus that is variable in size and shape.
This pelecypod burrows deeply, as much as 30 cm below the sediment- water
interface, and thus is often preserved intact, in living position. All records for the
genus in Ontario are for Mya arenaria and are from the Cornwall area. In Quebec the
genus is widely represented by M. arenaria and two other species. Most occurrences
are in sand.

SPECIES Mya arenaria Linne

references Chalmers ( 1907), Terasmae (1960, 1965), Wagner (1970), Whiteaves
(1901), Whittaker (1922)

Hiatella Bosc (Daudin ms) Figure 15C, D

The shell is highly variable in form because the animal nestles among pebbles,

Fig. 14 Macoma

Macoma balthica (Linne)

A Interior of right valve, x 3, GSC 551 18, Harrisons Corners, Stormont County,

Ontario.
B Exterior of left valve, x 3, GSC 55119, Harrisons Corners, Stormont County,

Ontario.
Macoma calcarea (Gmelin)

C Interior of left valve, x 1.5, GSC 55120, Cornwall area, Ontario.
D Interior and exterior view, x 1, GSC 20158, St-Maurice, Quebec. This specimen

shows detail of the pallial sinus more clearly.

cobbles, and even boulders, and often burrows into soft rocks. The shape may be
quadrate, trapezoidal, or irregular, with a posterior gape. Usually the shell is chalky
and heavy and has irregular growth striae. It may reach a length of 75 mm, although
most individuals are much less than that. Adults characteristically have an edentulous
hinge, but juveniles sometimes have a single, small, ephemeral cardinal tooth in the
right valve and two teeth in the left valve. The pallial line is discontinuous and the
sinus is small.

The one species encountered is the Saxicava arctica of earlier authors, the source
of the name for the Saxicava sand. Juveniles of this species have two rows of spines
running from the umbones to the posterior margin; these spines normally disappear
with age. This common species is often found in sand and gravel pits or other
exposures of these materials.

Mya Hiatella

Mya arenaria Linne

A Interior and exterior views of a right valve, x 1 , gsc 20160, St-Remi, Quebec.

B Interior of a broken left valve showing the chondrophore, x 3, GSC 55121,

Cornwall area. Ontario.
Hiatella arctica (Linne)

C Interior of right valve, x 3, GSC 55122, northwest of Moulinette, Ontario.
D Exterior of left valve, x 3, GSC 55123, northwest of Moulinette, Ontario^

ONTAR/G^

•<>

SPECIES Hiatella arctica (Linne) [Saxicava arctica and Saxicava rugosa are
synonyms]

REFERENCES Ami (1884, 1892, 1897, 1902, 1906), Antevs (1925, 1939), Chalmers
(1907), Coleman (1901a, 1901b, 1941), G. M. Dawson (1897), J. W. Dawson
(1857, 1871, 1883b, 1893), Goldring (1922), Johnston (1916, 1917), Keele and
Johnston (1913), Kindle (1918), Leidy (1856), Logan (1847, 1863), Lowdon and
Blake (1973, 1976, 1979), Murray (1852), Richards (1962), Terasmae (1960,
1965), Wagner (1958, 1970), Whiteaves (1901), Whittaker (1922), A. E. Wilson
(1956), W. J. Wilson (1898)

Conchostraca (Clam Shrimps) Figure 16

Conchostracans belong to the crustacean class Branchiopoda. The body is enclosed
within a translucent bivalve shell which may show various forms of ornamentation in
addition to growth lines. Genera are 2 to 16 mm in length and inhabit both freshwater
and marine environments.

Cyzicus Audoin Figure 16

This genus is characterized by a thin, pellucid, laterally compressed, subovate shell.
The umbonal area is small. There are numerous growth lines. Representatives of the
genus attain a length of about 12 mm.

Several incomplete specimens have been found in concretions from Green Creek,
near Ottawa. The whereabouts of this material is unknown, and so a diagrammatic
presentation of the genus is shown instead.

SPECIES Cyzicus dawsoni (Packard) [the original listing is Estheria dawsoni]
references Coleman (1932), J. W. Dawson (1893)

Fig. 16 Diagrammatic presentation of the genus Cyzicus, x3.

Cirripedia (Barnacles) Figures 17 and 18

Anyone who has visited the seacoast will be familiar with the myriads of barnacles
encrusting any suitable base of attachment from high-tide level down to the sea floor.

Complete fossils of this crustacean are rarely found. The usual remains are isolated
individual wall plates, opercular valves, or opercular plates. Less commonly wall
plates are attached to a calcareous basis (basal disc) and are intact. Often only the
basal disc will be found, adhering to a stone. Barnacle shell morphology is illustrated
in Figure 17.

Three species have been identified in Ontario, only one of which, Balanus
crenatus, is at all common. It is most abundant in the eastern part of the province, in

the Cornwall area. B. balanus has been reported only from the Ottawa district, andfi.
hameri , although more widely distributed, is also rare.

Balanus da Costa

The wall of Balanus is formed of six usually rigidly articulated plates. Externally the
wall plates may be either ribbed or merely rough. The basis is calcareous.

Balanus sensu stricto Figure 18 A -D

The central section (paries) of each wall plate has parietal tubes, whereas the radii
(overlapping flanges on either side of the paries) are solid. Species attain a length of
about 40 mm.

No complete fossil specimens were available, and so modern ones have been
substituted for illustration.

SPECIES Balanus (Balanus) balanus (Linne); Balanus (Balanus) crenatus Bruguiere
references Ami (1892, 1897, 1902, 1906), Antevs (1925, 1939), Coleman
(1901b), Goldring (1922), Kindle (1918), Terasmae (1960, 1965), Wagner (1958,
1970), Whittaker (1922), A. E. Wilson (1956), W. J. Wilson (1898)

Balanus (Chirona) Gray Figure 18E-G

In this subgenus both paries and radii are thin and solid. Specimens may reach a

length of about 65 mm.

SPECIES Balanus (Chirona) hameri (Ascanius)

references Ami (1906), Wagner (1970), Whittaker (1922)

ORIFICE

ALA

RADIUS

PARIES

Fig. 17

BASIS

Morphological terms applied to barnacles.

A 1. Rostrum, 2. Rostrolateral, 3. Lateral. 4. Carinolateral, 5. Carina.

B Compartment with ala on one side and radius on the other.

C 1. Scutum, 2. Tergum.

Isopoda (Isopods) Figure 19

These malacostracan crustaceans are often, but not always, shrimplike. The isopod
body is elongate, flattened, and made up of a cephalon with seven thoracic and six
abdominal somites. The last abdominal somite, or telson, is broad. A familiar
terrestrial member of this group is the sow-bug, or pill-bug, common under leaves or
piles of decaying vegetation. Other isopods are found in freshwater or marine
habitats.

Mesidotea Richardson Figure 19

This form attains a length of about 120 mm. The first thoracic somite envelopes the
cephalon laterally; the other thoracic somites are folded downwards laterally. The
eyes are dorsal. The length of the pleotelson is more than one quarter that of the body.
A single specimen was discovered in a concretion from the bank of the Ottawa
River east of Ottawa.
SPECIES Mesidotea sabini Kroyer
reference Kindle (1928)

Asteroidea (Starfish) Figure 20

These echinoderms have relatively broad, hollow arms that are not normally
separated from the central disc. The number of arms may vary from five to many. The
arms and disc have a skeleton of calcified ossicles. The ossicles frequently bear
spines or have granules distributed over the surface. There is an ambulacral groove on
the oral (downward-facing) side of each arm. Asteroids are free-moving. They are
capable of regenerating an arm where one has been thrown off to escape capture.
General features of asteroids are depicted in Figure 20A.

Crossaster Muller and Troschel Figure 20B

In this genus the arms, which are half as long as the breadth of the disc, may number
as many as 15. Individual specimens range between 200 and 300 mm in diameter. At
the points of union of the ossicles on the upper (aboral) surface there are club-shaped
tubercles that bear smaller spines, giving the arms a distinctive tufted appearance.

Starfish remains are not common. Specimens have been reported only from Green
Creek in Ontario and from Montreal in Quebec. No specimen of C. papposus was
available, and so a sketch has been prepared.

Fig. 18 Balanus (Balanus) Balanus (Chirona)
Balanus (Balanus) balanus (Linne)

A Top view of adult specimen, x 1, GSC 20190, Atlantic Coast.
B Side view of GSC 20190.
Balanus (Balanus) crenatus Bruguiere

C Top view of cluster of three individuals, x 1, GSC 20191, Atlantic Coast.
D Side view of GSC 20191.
Balanus (Chirona) hameri (Ascanius)
E Side view of adult specimen attached to a wall plate of another individual of B.

hameri, x 1, GSC 20192, St-Philomene, Quebec.
F An opercular plate (scutum), x 1, GSC 55124, Russell County, Ontario.
G A wall plate (or compartment), x 1, GSC 55125, Russell County, Ontario.

Fig. 19 Mesidotea

Mesidotea sabini Kroyer
Specimen preserved in a
concretion, x 1, gsc 9368,
Ottawa River, east of Ottawa,
Ontario

SPECIES Crossaster papposus (Linne)

REFERENCES Dawson (1893), Goldring (1922), Whiteaves (1901)

Ophiuroidea (Brittlestars) Figures 21 and 22

Ophiuroids are related to the starfish but are distinct in that the arms and central disc
are strongly differentiated. Also, the arms are slender and elongate. An individual
may throw off pieces of an arm or may completely dismember itself without
permanent injury; the lost parts are readily reproduced. Figure 21 illustrates a typical
brittlestar.

Four genera have been identified from the Champlain Sea, only two of which have
been found in Ontario. Isolated ossicles are sometimes found in the silts and clays;
more complete specimens are much rarer.

AMBULACRAL GROOVE

Fig. 20 Asteroidea Crossaster

A Morphological terms applied to starfish (oral view).

Crossaster papposus (Linne)

B Schematic presentation of a typical specimen (aboral view).

VENTRAL SHIELD

BUCCAL SHIELD
Fig. 21 Morphological terms applied to brittlestars (oral view).

MOUTH

t miu ffii i. i. m,«m.mimmMmm*iimmim

«'■' /Mi t'/^t-Xr, ...

J 1 > ,- fll

Fig. 22 Ophiura

Ophiura sarsi Liitken

Impression in clay, x 3, Ottawa, Ontario.

Ophiocoma Agassiz

This genus has comparatively stout arms that are widest at a distance away from the
base. The arm spines are long and solid and the disc is granulate.

Specimens have been reported from several Champlain Sea localities, from Quebec
City in the east to Ottawa in the west, but none was identifiable as to species.
SPECIES Ophiocoma sp.
references Dawson (1871, 1893)

Ophiura Lamarck Figure 22

The central disc in this genus is covered with scales. Rudimentary dorsal arm plates
fill the notches around the edge of the disc at the bases of the arms. Ventral shields on
the arms are usually triangular and are more broad than long. Spines on the arms are
vestigial.

The genus has been identified from Montreal and from the Ottawa area; specimens
are rare.

SPECIES Ophiura sarsi Liitken; Ophiura sp.
references J. W. Dawson (1857), Wagner (1954)

Polychaeta (Polychaete Worms) Figure 23

The Annelida, the phylum to which the polychaete worms belong, have a distinct
head, a segmented trunk, and an unsegmented pygidium. In the class Polychaeta the
trunk segments have lateral bundles of bristles termed chaetae. The chitinous jaws are
distinctive and are often the only part to be fossilized, although the burrows and
calcareous tubes formed by some families are also preserved. Most of these worms
are marine, with a comparatively small number of representatives living in fresh or
brackish waters.

Two of the three genera recognized in the Champlain Sea deposits have been
reported from Ontario.

Nereis Linne Figure 23A

Some species may attain a length of 450 mm or more; the species N. pelagica,
recorded only from Green Creek, reaches a maximum of about 125 mm for females
and 50 mm for males. The first segment of the body usually has four tentacles on
each side, and the last segment has a pair of long cirri that give the impression of a
divided tail.

It has been necessary to provide a sketch for this genus.
SPECIES Nereis pelagica Linne
REFERENCES Dawson (1891, 1893), Goldring (1922), Whiteaves (1901)

Serpula Linne Figure 23B

This is a tube-building polychaete. The irregularly tapering calcareous tubes are
coiled and contorted and have fine concentric ridges on the surfaces. They are
attached at the lower end, usually to a shell or rock, and are more or less erect above
their point of attachment. The tubes may reach a length of about 100 mm.

Only two localities are known for this genus in Ontario, both in the Ottawa district.
Neither specimen was available for illustration, and therefore a drawing has been
substituted.

SPECIES Serpula vermicularis Linne
references Goldring (1922), Wagner (1970)

Fig. 23 Nereis Serpula

Nereis pelagica Linne

A Diagrammatic presentation, x 1.

Serpula vermicularis Linne

B Diagrammatic presentation of several calcareous tubes, X 0.7.

Vertebrata

Chordata Figures 24 to 32

A vertebrate find is an exciting experience. The most rewarding locale is the area
between Green Creek and Hiawatha Park on the south side of the Ottawa River,
where concretions bearing relatively complete fish skeletons are commonly eroded
out of the clays. The concretions have yielded six genera of fish and, as well, some
limb bones and part of the lower jaw of a young seal. Seal and whale bones are rare in
the Champlain Sea; they have been found in sand and gravel deposits near Ottawa.

Pisces

Figure 24 illustrates various terms related to fish morphology.

Clupeiformes

This is the order of fish to which the capelin and smelts belong. They have a short
dorsal fin, the base of which is shorter than one-third of the total length of the body
(the total body length is the distance from the tip of the snout to the end of the
vertebral column). The dorsal fin is situated at about the midpoint of the body, and
the base of the pelvic fin is below the dorsal fin.

Mallotus Cuvier Figure 25

Capelin may grow as long as about 230 mm but are usually shorter. The body is
elongate and compressed. The length of the head (the distance from the tip of the
snout to the most posterior part of the opercular membrane) is about one-quarter of
the total length. The dorsal fin, inserted about midway between the snout and the tip
of the tail, has 12 to 15 rays. The anal fin, with 18 to 23 rays, has a longer base than
that of the dorsal fin. The two pelvic fins are inserted below the dorsal fin, and the

NOSTRIL
EYE

1ST DORSAL FIN 2ND DORSAL FIN CAUDAL FIN
■RAY

OPERCULAR MEMBRANE
(GILL COVER)

Fig. 24 Morphological terms applied to fish.

CAUDAL PEDUNCLE
ANAL FIN

pectoral fins are just behind the gill openings. The pectoral fins are broadly based.

The scales on this genus are small.
Specimens have been collected from the Ottawa River drainage area above Ottawa,

as well as from the classic Green Creek locality.

SPECIES Mallotus villosus (Muller)

REFERENCES Ami (1884, 1887, 1897, 1902), Champagne, Harington, and
McAllister (1979), Coleman (1901b), J. W. Dawson (1871, 1878, 1891, 1893),
Ells (1907), Goldring (1922), Harington (1971, 1972, 1977), Johnston (1917),
Lambe (1914), Leidy (1856), Logan (1863), Lyell (1845), McAllister, Cumbaa,
and Harington (1981), Murray (1852), Wagner (1970)

Osmerus Lacepede Figure 26

Smelts are slender fish; their body is deepest anterior to the dorsal fin and tapers
towards the head and tail. The head is pointed, with the lower jaw projecting slightly
beyond the upper. Individuals may reach a length of 350 mm, but most range
between 200 and 250 mm in length. The dorsal fin, with 9 to 1 1 rays, is situated at
about the middle of the back; this fin is higher than long. The anal fin has 15 to 18
rays and is longer than it is high. The caudal fin is deeply forked. Pectoral fins lie
immediately behind the gill openings, and the abdominally placed pelvic fins are
below the anterior part of the dorsal fin.

Like Mallotus, Osmerus has been found in a concretion from Green Creek. In
Osmerus the mouth opening extends behind the eyes, whereas in Mallotus the mouth
extends posteriorly only to about the middle of the eye. There are also differences in
the numbers of the rays in the dorsal and anal fins and in body proportions.
SPECIES Osmerus mordax (Mitchill)
references Champagne, Harington, and McAllister (1979), Coleman (1901b),

J. W. Dawson (1891, 1893), Harington (1971, 1972, 1977), McAllister, Cumbaa,

and Harington (1981)

Gadiformes

Specimens of this order from Ontario are of the tomcod (genus Microgadus).

Microgadus Gill Figure 27A

The tomcod is slender, reaching a maximum length of about 300 mm. The mouth is
short and there is a small barbel hanging from the lower jaw. This genus differs from
most in having three dorsal fins rather than the usual two. The caudal fin is rounded.
The anal fin is divided into two parts. Pectoral fins reach backwards as far as the vent,
but the pelvic fins below them are shorter.

Two specimens were found in concretions east of Ottawa in 1979. A sketch
illustrating a whole fish has been substituted for the line drawing of part of an
individual that accompanied the report of the presence of this genus in Ontario.
SPECIES Microgadus tomcod Walbaum
reference McAllister, Cumbaa, and Harington (1981)

Gasterosteiformes

Fish belonging to this order are usually small, 150 mm or less in length. The dorsal,
anal, and pelvic fins have sharp spines. Sticklebacks are members of this order.

Fig. 25 M allot us

Mallotus villosus (Miiller)
Specimen in a concretion, x 1,
GSC 6597a, Green Creek, Ontario

Fig. 26 Osmerus

Osmerus mordax (Mitchill)
Skeleton preserved in a concretion,
x 1 . 1 , NMC 3580 1 B , Green Creek
area, Ontario.

Gasterosteus Linne Figure 27B

Sticklebacks have a rather stout body with a very slim caudal peduncle. Maximum
length is 100 mm, but most individuals do not exceed about 75 mm. The lower jaw
of the small mouth protrudes beyond the upper. A characteristic feature is the
presence of three, on rare occasion four, isolated serrated spines in front of the dorsal
fin. The dorsal fin has 10 to 14 (usually 12) rays. The caudal fin is truncate, not
forked. Origin of the anal fin is behind that of the dorsal. The pelvic fin has one soft
ray plus a single strong spine, and the pectoral fins are large and are situated a short
distance behind the gill opening.

The concretions of Green Creek and vicinity have provided the only remains of this
fish known in the Champlain Sea deposits.
SPECIES Gasterosteus aculeatus Linne; Gasterosteus sp.

REFERENCES Ami (1902), Champagne, Harington, and McAllister (1979), Coleman
(1901b), J. W. Dawson (1871, 1878, 1891, 1893), Goldring (1922), Harington
(1971, 1972, 1977), McAllister, Cumbaa, and Harington (1981)

Perciformes

Two of the fish identified from the Champlain Sea, the sculpins (Artediellus) and
lumpsuckers (Cyclopterus), belong to this large order.

Artediellus Jordan

The body is elongate but stout. Sculpins are generally small, and the possible species
from the concretions does not reach more than 100 mm in length. The large head,
with prominent eyes placed high, is distinguished by a long, hooklike spine on each
cheek. The gill coverings end in a covered spine high on each side. The mouth is
terminal and ends under the middle of the eye. Two dorsal fins are present: a soft
anterior one beginning over the gill opening, followed by a spinous one with 12 to 14
rays. The caudal fin is small and spade-shaped. The anal fin, with 10 to 12 rays, is
situated under the second dorsal fin. The pectoral fins are large, reach back as far as
the beginning of the anal fin, and have 20 to 22 rays. The long, slender pelvic fins are
on the ventral edge under the forward edge of the pectorals.

In the Champlain Sea deposits Artediellus has been found only in the Green Creek
area. This specimen should properly be referred to as Artediellus sp. until it can be
confirmed whether it is A. uncinatus or A. atlanticus. The whereabouts of the
specimen is unknown and no illustration was available.
SPECIES Artediellus sp. [recorded previously as Cottus sp., Cottus uncinatus, Cottus

(Centrodermichthys) uncinatus, and Artediellus uncinatus (Reinhardt)]
REFERENCES Ami (1897, 1902), Champagne, Harington, and McAllister (1979),

Coleman (1901b), J. W. Dawson (1878, 1891, 1893), Harington (1971, 1972,

1977), Johnston (1917), Logan (1863), McAllister, Cumbaa, and Harington

(1981)

Cyclopterus Linne

The lumpsucker has a stout, thick body with a small head. The eye is small, as is the
mouth; the angle of the mouth is in front of the eye. Maximum length for specimens
of the species from the Champlain Sea, as recorded by J. W. Dawson (1871), is
610 mm, but specimens found in concretions would be much smaller than this. The
anterior dorsal fin is visible only in small specimens (less than 30 mm long); in larger
specimens it is enclosed in a hump that is covered in hard, wartlike tubercles. The

Fig. 27 Microgadus Gasterosteus

Microgadus tomcod Linne

A Sketch to show general features of the species.

Gasterosteus aculeatus Linne

B Drawing of a specimen in a concretion, from J. W. Dawson (1893), x 0.6.

posterior dorsal fin has 10 or 1 1 rays and is located behind the hump. The caudal fin is
slightly rounded. The anal fin, with 9 to 1 1 rays, is below the posterior dorsal fin and
is similar in size to it. Pectoral fins are large and extend along the lower two-thirds of
the gill opening; they have 20 or 21 rays. Pelvic fins, situated on the ventral side
between the bases of the pectorals, are reduced to fleshy knobs, which, with a
surrounding circular flap of skin, form sucking discs, a feature not likely to be
preserved in fossil specimens.

Like the other genera of fish, Cyclopterus has been identified from a concretion
from Green Creek. The whereabouts of this specimen is unknown and no illustration
was available.

SPECIES Cyclopterus lumpus Linne [one early record shows it as Cyclopteris]
references Ami (1884, 1897), Champagne, Harington, and McAllister (1979),

Coleman (1901b), Dawson (1871, 1878, 1891, 1893), Goldring (1922), Harington

(1971, 1972, 1977), Leidy (1856), Logan (1863), McAllister, Cumbaa, and

Harington (1981), Murray (1852)

Mammalia

Two orders of marine mammals, namely Cetacea (whales) and Pinnipedia (seals), are
represented. Most finds have been of isolated bones or incomplete skeletons,

although essentially complete seal skeletons were found at Montreal and Tetreauville,
Quebec, and whale skeletons were unearthed at Cornwall, Ontario, and Daveluyville,
Quebec.

Identification of bones would need the services of an expert, and I will not attempt
to describe whale and seal skeletons here. The larger museums [Royal Ontario
Museum (ROM) in Toronto and the National Museum of Natural History (nmc) in
Ottawa] have vertebrate palaeontologists on staff, and vertebrate specialists may be
contacted in some of the universities in the province.

Cetacea

Delphinapterus Lacepede Figure 28

This is the white whale, or beluga, a small whale that may reach a length of 5 m but
that usually has a maximum length between 3 and 4 m. White whales are common on
the Gulf of St. Lawrence at present and often travel up the St. Lawrence River as far
as Quebec City. Bones have been found in the Champlain Sea sands in the vicinity of
Ottawa (Jock River, Ottawa East, Rideau Junction, Uplands) and at Pakenham,
Cornwall, and Williamstown.
SPECIES Delphinapterus leucas Pallas [recorded as Beluga catodon. Beluga

vermontana, Delphinapterus catodon, Delphinapterus {Beluga) catodon, and

Delphinapterus vermontanus ]
references Billings (1870), Coleman (1901a), J. W. Dawson (1871, 1893), Dyck

et al. (1966), Harington (1971, 1972, 1977, 1981), Lambe (1910, 1914),

Laverdiere (1950), Lowdon and Blake (1979), Perkins (1908), Selwyn (1872),

Sternberg (1951), Whiteaves (1907)

Megaptera Gray

Part of a skeleton of a humpback whale was found in a gravel pit at an elevation of
about 128 m (420 ft) at Warwick, near Smiths Falls, Ontario, in 1882. The female
humpback whale averages about 15 m in length; males are smaller.
SPECIES Megaptera novaeangliae Borowski [M. longimana Gray is a synonym]
REFERENCES Coleman (1901a, 1901b, 1941), J. W. Dawson (1883a, 1883b, 1893),
Harington (1971, 1972, 1977, 1981), Laverdiere (1950)

Balaena Linne Figure 29

A characteristic of the bowhead whale is that the length of its head is more than

one-third that of its body. Total length is about 20 m. The baleen plates, of which

there are approximately 360 on each side of the mouth, measure 35 cm wide by 3 m

long. Bones of a right pectoral flipper, ribs, and part of a jaw, all probably from one

individual, were collected near White Lake, Ontario.

SPECIES Balaena mysticetus Linne

REFERENCES Harington (1977, 1981), Lowdon and Blake (1979)

Pinnipedia

Phoca Linne

This is a genus of aquatic mammals in which the limbs have developed as flippers.
Generally the females are much smaller than the males. Two subgenera are
represented in the Champlain Sea.

„-«!*£

Fig. 28 Delphinapterus

Delphinapterus leucas Pallas

Cranium (top view showing blow hole) and mandibles, x 0.3, NMC 21336, near

Pakenham, Ontario.

Fig. 29 Balaena

Balaena mysticetus Linne

Right humerus, x 0.3, NMC 29414, near White Lake, Renfrew area, Ontario.

Phoca (Pagophilus) Gray Figure 30

Harp seals attain a maximum length of just under 2 m. Isolated bones that may be of
this genus have been reported from the clays in and around Ottawa. The only
specimen that can be referred definitely to a harp seal is part of the lower jaw of a
young individual preserved in a concretion at Green Creek.
SPECIES Phoca (Pagophilus) groenlandica (Erxleben) [listed in earlier records as

Phoca groenlandica, Pagophilus groenlandicus , and Phoca sp.]
references Ami (1884, 1892, 1897, 1902), Coleman (1910b), J. W. Dawson

(1878, 1893), Grant (1883), Harington (1971, 1972, 1977, 1981), Laverdiere

(1950), Sternberg (1951)

Phoca (Pusa) Scopoli Figure 31

The ringed seal is the smallest of the pinnipeds, with an average length of 1.4 m. At
present, it is seldom found south of the Strait of Belle Isle. A single bone was found
in 1975, in a sand pit near Uplands (now Ottawa International) Airport, Ottawa.
However, the almost complete specimen from Tetreauville (now in the western part
of Hull), Quebec, is much more impressive and has therefore been used to illustrate
this form.

SPECIES Phoca hispida Schreber
reference Harington (1977, 1981)

Erignathus Gill Figure 32

Bearded seals reach a maximum length of just under 2 m when adult. These seals
usually feed in shallow water, and molluscs form an important part of their diet. Part
of a cranium, minus teeth, was collected near Finch, Ontario.
SPECIES Erignathus barbatus Erxleben
references Harington (1977, 1981)

Fig. 30 Phoca (Pagophilus)

Phoca (Pagophilus) groenlandica (Erxleben)

Drawing of a part of the lower jaw of a young harp seal in a concretion from Green

Creek, from J. W. Dawson (1893).

Fig. 31 Phoca (Pusa)

Phoca (Pusa) hispida Schreber

Ringed seal skeleton (NMC 6830) on display in the National Museum of Natural
Sciences, Ottawa. Specimen from Tetreauville, in the western part of Hull, Quebec.
NMC Photo 72-1540.

Fig. 32 Erignathus

Erignathus barbatus Erxleben

View of top of skull, x 0.8, ROM 985, near Finch, Ontario. NMC Photo 75-5772.

Nonmarine Species

Figure 33

Concretions from the Green Creek deposits have yielded a number of species of
nonmarine origin (both fauna and flora), suggesting that, at this point in time, the area
formed the margin of the retreating Champlain Sea. Among the species identified are
insects, freshwater fish, a small terrestrial mammal, birds (several feathers
discovered, Fig. 33A), and a variety of plants (Fig. 33B). Although these are not
species that owed their presence to the Champlain Sea, they are ones that apparently
had become fossilized in the marginal environment of the sea. The most
comprehensive list is given by Ami (1902). Harington (1971, 1972, 1978) and
Harington and Occhietti (1980) mention the more recent discoveries.

A paper by Gadd (1980) gives an up-to-date discussion of the conglomeration of
marine, freshwater, and terrestrial species found in the concretions at Green Creek
and vicinity, and of their possible relative ages.

Insecta

Trichoptera (caddisflies)
Phryganea ejecta Scudder Ami (1902), Coleman (1932)

Coleoptera (beetles)

Tenebrio calculensis Scudder Ami (1902), Coleman (1932), Scudder (1895)
Fornax ledensis Scudder Ami (1902), Coleman (1932), J. W. Dawson (1893),
Scudder (1895)
Byrrhus ottawensis Scudder Ami (1902), Coleman (1932), Scudder (1895)

Pisces (freshwater fish)
Myoxocephalus thompsoni (Girard) [deepwater sculpin] Champagne, Harington,
and McAllister (1979), McAllister, Cumbaa, and Harington (1981)
Salvelinus {Cristivomer) namaycush (Walbaum) [lake char] Champagne,
Harington, and McAllister (1979), Harington (1971, 1972, 1978), McAllister,
Cumbaa, and Harington (1981)

Coregonus cf. C. artedii Le Sueur [cisco] McAllister, Cumbaa, and Harington
(1981)

Mammalia

Martes americana (Turton) [marten] Harington (1971, 1972)

Aves

Feathers of "small wading birds", according to J. W. Dawson, plus a single bird
vertebra from the sands near Uplands (now Ottawa International) Airport, Ontario.
J. W. Dawson (1893), Harington (1971, 1972, 1978), Harington and Occhietti
(1980).

Fig. 33 Nonmarine fossils

A Impression of a feather in a concretion, x 1, GSC 6600, Green Creek, Ontario.
B Leaf impression in a concretion, x 1, GSC 32381, Green Creek, Ontario.

Plantae (listed alphabetically)

Ami (1902), J. W. Dawson (1893), Penhallow (1900)

Descriptions and illustrations of species for which common names are given can be

found in McKay and Catling (1979) or Niering and Olmstead (1979). Names, and

therefore identity, of the other plants could not be confirmed.
Acer saccharinum (silver maple)
Acer spicata (mountain maple)
Algae spp.
Alnus sp. (alder)

Arctostaphvlos uva-ursi (bearberry)

Betula lutea (yellow birch — now known as B. alleghaniensis)
Brasenia peltata
Bromus ciliatus ?
Carex magellanica
Carices or Gramineae spp.
Cyperaceae (sedges)

Drosera rotundifolia (round-leaved sundew)

Elodea canadensis [=Anacharis canadensis] (common waterweed)
Encyonema prostratum ?
Equisetum limosum ?
Equisetum scirpoides ?
Equisetum sylvaticum (woodland horsetail)
Fontinalis sp.
Fucus digitatus
Fucus sp. or Ulva sp.

Gaylussacia resinosa [=G. baccata] (black huckleberry)
Gaylussacia sp. (huckleberry)
Hypnum fluitans
Graminea resinosa ?
Oryzopsis asperifolia
Populus balsamifera (balsam poplar)
Populus grandidentata (large-toothed aspen)
Potamogeton natans
Potamogeton pectinatus ?
Potamogeton perfoliatus ?
Potamogeton pusillus ?
Potamogeton rutilans
Potentilla anserina (silverweed)
Potentilla canadensis (Canadian dwarf cinquefoil)
Potentilla norvegica (rough cinquefoil)
Potentilla tridentata
Thuja occidentalis (eastern white cedar)
Trifolium repens (white clover)
Typha latifolia (common cattail)
Valisneria spiralis ?
Valisneria sp. (tape grass)

Repositories of Illustrated Specimens

GSC Geological Survey of Canada
NMC National Museums of Canada
ROM Royal Ontario Museum

Glossary of Morphological Terms

abaxial (Gastropoda) — outwards away from the shell axis.

adapical (Gastropoda) — towards the apex of the shell, along the axis or slightly

oblique to the axis.

adaxial (Gastropoda) — inwards towards the shell axis.

adductor muscle (Pelecypoda) — a muscle, commonly one of two, connecting the

two valves and tending to draw them together.

ambulacral groove (Asteroidea) — an axial depression along the oral surface of the

arm and covered by a series of ambulacral ossicles.

ambulacrum (Asteroidea) — one of the radial areas bearing the tubular protrusion by

which locomotion is accomplished.

anal fin (Pisces) — a median unpaired fin on the ventral margin between the anus and

the caudal fin.

anterior canal (Gastropoda) — see siphonal canal.

anteroposterior (Pelecypoda) — in a front-to-back direction.

aperture (Gastropoda) — an opening at the last-formed margin of the shell, providing

an outlet for the head and foot.

apex (Gastropoda) — top of the shell.

apical (Gastropoda) — referring to the apex.

aster (Porifera) — microsclere rays diverging from a central point.

avicularium (Bryozoa) — a specialized zooid with smaller polypide but with strong

muscles that operate a mandiblelike operculum; resembles the head of a bird.

axial (Gastropoda) — parallel or subparallel with the axis of the shell.

axis (Gastropoda) — an imaginary line through the apex of the shell and about which

the whorls are coiled.

baleen (Cetacea) — one of the fibrous plates in the mouth through which food is

strained.

barbel (Pisces) — a slender, whiskerlike tendril on the head of certain fishes.

basis; pi. bases (Cirripedia) — a calcareous or membraneous plate serving to attach

the barnacle to the substrate or to a foreign body.

beak (Pelecypoda) — a noselike angle located along or above the hinge margin and

marking the point where growth of the shell started.

calcareous (general) — composed of calcium carbonate.

callus (Gastropoda) — a smooth, shelly layer on the parietal region or extending from

the inner lip over the base or into the umbilicus.

cardelle (Bryozoa) — same as condyle.

cardinal tooth (Pelecypoda) — a hinge tooth situated close to the beak.

carina (Cirripedia) — a single compartment plate situated at the end of the shell where

the cirri are protruded.

caudal fin (Pisces) — a terminal vertical fin; tail.

caudal peduncle (Pisces) — a slender posterior portion of the body situated between

the anal and caudal fins.

centrum (Porifera) — the middle part of a spicule from which rays diverge irregularly.

cephalon (Isopoda) — head.

chaeta; pi. chaetae (Polychaeta) — a bristle.

chamber (Porifera) — a cavity containing the operative flagellate cells.

chitinous (Polychaeta) — horny.

chondrophore (Pelecypoda) — a process with a hollowed-out surface for attachment

of the internal ligament.

cirrus; pi. cirri (Polychaeta) — a filament.

collabral (Gastropoda) — conforming to the shape of the outer lip, as shown by

growth lines.

columella (Gastropoda) — a solid or hollow pillar surrounding the axis of a coiled

shell and formed by the adaxial walls of the whorls.

concentric (Pelecypoda) — having a direction coinciding with that of the growth

lines.

condyle (Bryozoa) — a rounded protuberance for hingement of the operculum.

corneous (Gastropoda) — consisting of a horny substance.

crenate (Pelecypoda) — with notches along the edge or crest (as of ribs).

crenulate (Pelecypoda) — minutely crenate.

disc (Asteroidea, Ophiuroidea) — the central part of the body, more or less distinctly

separable from the arms.

dorsal (Pelecypoda) — referring to the region of the hinge.

dorsal fin (Pisces) — a fin developed on the back.

edentulous (Pelecypoda) — lacking hinge teeth.

euaster (Porifera) — an aster lacking a centrum.

foliate (Bryozoa) — of thin, leaflike layers.

fusiform (Gastropoda) — a slender spindle shape tapering almost equally towards

both ends.

globose (Gastropoda) — more or less spherical, rounded.

groove (Asteroidea)— see ambulacra! groove.

growth line (Pelecypoda, Gastropoda) — a line on the surface of the shell marking the

position of the shell margin at some stage of growth.

helicocone (Gastropoda) — a distally expanding coiled tube that forms most gastropod

shells.

hinge plate (Pelecypoda) — a shelly internal platform bearing hinge teeth and situated

below the beak, adjacent to parts of the dorsal margin.

hinge tooth (Pelecypoda) — a shelly structure, usually one of a series, adjacent to the

dorsal margin, and received by a socket in the opposite valve; hinge teeth serve to

hold the closed valves in position.

imperforate (Bryozoa) — lacking perforations.

infaunal (Pelecypoda) — living burrowed into the substrate.

inner lip (Gastropoda) — the adaxial margin of the shell aperture, extending from the

foot of the columella to the suture.

involute (Gastropoda) — having the last whorl enveloping the earlier ones, so that the

height of the aperture corresponds to the height of the shell.

ligament (Pelecypoda) — a horny, elastic structure, or structures, joining the two

valves of the shell dorsally and serving as a spring, causing the valves to open when

the adductor muscles relax.

longitudinal septum (Cirripedia) — a wall of tubes disposed at right angles to the

inner and outer laminae of the compartment plate and separating them.

lunule (Pelecypoda) — a depression, commonly heart-shaped, situated anterior to the

beaks in many bivalves.

megasclere (Porifera) — a spicule that is an element of the framework of the

organism.

microsclere (Porifera) — a spicule that is loose in the flesh and does not form a part of

the skeletal framework; microscleres are rare in fossils.

nacreous (Gastropoda, Pelecypoda)— lustrous, pearly.

opercular membrane (Pisces) — a gill cover.

opercular valve (Cirripedia) — a moveable plate in the orifice.

operculum (Bryozoa) — a small calcareous or chitinous lamina articulating on

condyles (cardelles) that project from the edge of the orifice.

operculum (Gastropoda) — a corneous or calcareous structure borne by the foot and

serving for closure, wholly or in part, of the aperture.

opisthogyrate (Pelecypoda) — curved so that the beaks point posteriorly; a term

applied to the umbones.

oral (Asteroidea, Ophiuroidea) — referring to the surface of the animal that contains

the mouth and that is directed downwards.

orifice (Bryozoa) — the primary opening of the zooecium, for extrusion of the

polypide.

orifice (Cirripedia) — an opening in the upper part of the barnacle shell, containing the

opercular valves.

ossicle (Asteroidea, Ophiuroidea) — any individual calcified element of the skeleton,

usually referring to the larger of such elements.

outer lip (Gastropoda) — abaxial margin of the aperture, extending from the suture to

the foot of the columella.

ovate (Pelecypoda) — shaped like the longitudinal section of an egg.

ovicell (Bryozoa) — any structure serving to contain the larvae during their

development.

pallial line (Pelecypoda) — a line or narrow band on the interior of the valve, close to

the margin, and marking the line of attachment of the marginal muscles of the mantle.

pallial sinus (Pelecypoda) — an embayment of the pallial line, forming the line of

attachment of the siphonal retractor muscle.

paries; pi. parietes (Cirripedia) — the median, triangular part of the compartment

(wall) plate, with the lower edge attached to the basis.

parietal region (Gastropoda) — an area of the basal surface of the helicocone, just

inside and just outside the aperture.

parietal tube (Cirripedia) — one of the myriad porelike canals in the longitudinal

septum.

pectoral fin (Pisces) — either of a pair of fins usually situated behind the head, one on

each side; pectoral fins correspond to the forelimbs of higher vertebrates.

pellucid (Conchostraca) — translucent .

pelvic fin (Pisces) — either of a pair of fins on the lower surface of the body; these fins

correspond to the hind limbs of higher vertebrates.

perforate (Gastropoda) — with a cavity or depression.

periostracum (Pelecypoda) — a thin coat of horny material that covers the exterior of

the calcareous shell.

peristome (Gastropoda) — the margin of the aperture.

pleotelson (Isopoda)— a structure formed by the fusion of one or more abdominal

somites with the telson.

polypide (Bryozoa) — the soft parts of the zooid.

porcellaneous (Pelecypoda) — of translucent, porcelainlike appearance.

pseudocardinal (Pelecypoda) — an irregularly formed tooth situated close to the

beak.

punctations (Gastropoda) — minute pits.

pygidium (Polychaetia) — tail section.

quadrate (Pelecypoda) — square, or almost so.

radial (Pelecypoda) — a direction of growth outwards from the beak, commonly

indicated by the direction of ornamentaiton.

radiate (Porifera) — with spicules radiating outwards from a central point.

radius; pi. radii (Cirripedia) — the lateral part of a compartment plate adjoining the

paries and marked off from it by a change in the direction of growth lines and by a

depressed exterior surface.

resilifer (Pelecypoda) — a recess or process for the attachment of an internal ligament.

reticulate (Gastropoda) — ornamentation consisting of a network of obliquely

intersecting linear ridges.

rib (Gastropoda) — a round-topped elevation of moderate width and prominence,

disposed collabrally on the shell surface.

rostrate (Pelecypoda) — having a pointed, beaklike posterior end.

shoulder (Gastropoda) — angulation on the whorl forming the abaxial edge of the

sutural ramp or shelf.

siphonal canal (Gastropoda) — a tubular or troughlike extension of the anterior

(abapical) part of the apertural margin.

somite (Isopoda) — a division of the body in the cephalon, thorax, and abdomen; the

exoskeleton of each somite comprises a body-ring that is generally divisible into a

dorsal and a ventral portion.

sphaeraster (Porifera) — an aster with a globular centrum.

spicule (Porifera) — a unit of the mineral skeleton of a sponge.

spiral (Gastropoda) — as applied to ornamentation, passing continuously round the

whorls almost parallel with the suture.

spire (Gastropoda) — the adapical, visible part of all the whorls exclusive of the last

(or body) whorl.

spongin (Porifera) — an organic, tough, flexible material related chemically to horn

and hair.

stria; pi. striae (Pelecypoda) — a narrow linear furrow or raised line on the surface of

the shell.

striate (Gastropoda) — with a narrowly incised shallow groove, or grooves.

strongyle (Porifera) — a type of megasclere with a single axis and with both ends

bluntly rounded.

subglobose (Gastropoda) — almost rounded.

substrate — the base or material on, or in, which an organism lives.

sutural shelf (Gastropoda) — a horizontal flattened band in some shells that adjoins

the adapical suture of the whorls.

suture (Gastropoda) — a continuous line on the shell surface, where the whorls

adjoin.

taxodont (Pelecypoda) — with numerous, short hinge teeth, some or all of which are

transverse to the hinge margin.

telson (Isopoda) — the last somite of the body; tail.

trapezoidal (Pelecypoda) — with four straight sides, no two of which are parallel.

trigonal (Pelecypoda) — three-sided.

trochiform (Gastropoda) — with a flat-sided conical spire and an almost flat base.

umbilicus (Gastropoda) — a cavity or depression formed round the shell axis between

the faces of the adaxial walls of the whorls where these do not coalesce to form a solid

columella.

umbo (Conchostraca) — the apical portion of either valve.

umbo; pi. umbones (Pelecypoda) — the region of the valve surrounding the point of

maximum curvature of the longitudinal dorsal profile and extending to the beak when

not coinciding with it.

valve (Pelecypoda) — one of the calcareous structures (two in most pelecypods) of

which the shell consists.

ventral shield (Ophiuroidea) — an ossicle of secondary origin on the oral side of the

arm.

whorl (Gastropoda) — any complete coil of the helicocone.

zoarium (Bryozoa) — an assemblage of many zooids forming a bryozoan colony.

zooecium (Bryozoa) — a chitinous, double-walled sac, chamber, or tube containing

the soft parts or polypide.

zooid (Bryozoa) — a single bryozoan animal, consisting of soft parts and a skeleton.

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ISBN 0-88854-295-X
ISSN 0082-5093

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